Calls

• Moans, throat clatter, departure honk of male from nest
  
  Recorded by Gerrit Vyn
  Alaska June 2007
• Throat clatter and departure honk of male from nest
  
  Recorded by Gerrit Vyn
  Alaska June 2007
• Departure honk and arrival call of males at nests
  
  Recorded by Gerrit Vyn
  Alaska June 2007
• Moans at nest and throat clatter
  
  Recorded by Gerrit Vyn
  Alaska June 2007
• Moans, honks, and arrival calls of male at nest colony
  
  Recorded by Gerrit Vyn
  Alaska June 2007
• Arrival calls of male at nest
  
  Recorded by Gerrit Vyn
  Alaska June 2007
• Throat clatter and departure honk of male from nest
  
  Recorded by Gerrit Vyn
  Alaska June 2007
Courtesy of Macaulay Library
© Cornell Lab of Ornithology.

Low groans, croaks, or hisses.